Physicochemical properties GG GA GC GU AG AA AC AU CG CA CC CU UG UA UC UU Twist 0.347 0.347 2.2 0.347 -0.888 -0.27 0.347 0.965 -2.741 -0.27 0.347 -0.888 -0.27 0.347 0.347 -0.27 Tilt -0.211 1.321 -0.67 0.555 0.096 -1.896 0.555 1.015 -0.823 0.862 -0.211 0.096 0.862 -0.977 1.321 -1.896 Roll 1.652 0.413 -1.102 -1.698 0 -0.689 -1.698 -0.643 1.652 0.643 0.092 0 0.643 1.01 0.413 -0.689 Shift -0.551 0.147 0.147 1.545 -0.813 -1.163 1.545 -0.988 2.156 0.497 -0.551 -0.813 0.497 -0.639 0.147 -1.163 Slide -1.407 -0.969 0.729 0.51 0.127 1.386 0.51 0.893 -2.009 0.346 -1.407 0.127 0.346 0.401 -0.969 1.386 Rise 0.802 1.515 -0.386 -0.149 0.564 -0.862 -0.149 -0.149 0.564 -1.931 0.802 0.564 -1.931 0.089 1.515 -0.862